*,*::before,*::after{box-sizing:border-box;margin:0;padding:0}

:root{
    --color-canvas:#fffaf0;
    --color-surface-soft:#faf5e8;
    --color-surface-card:#f5f0e0;
    --color-surface-strong:#ebe6d6;
    --color-surface-dark:#0a1a1a;
    --color-surface-dark-elevated:#1a2a2a;
    --color-ink:#0a0a0a;
    --color-body:#3a3a3a;
    --color-body-strong:#1a1a1a;
    --color-muted:#6a6a6a;
    --color-muted-soft:#9a9a9a;
    --color-hairline:#e5e5e5;
    --color-hairline-soft:#f0f0f0;
    --color-primary:#0a0a0a;
    --color-primary-active:#1f1f1f;
    --color-on-primary:#ffffff;
    --color-on-dark:#ffffff;
    --color-brand-pink:#ff4d8b;
    --color-brand-teal:#1a3a3a;
    --color-brand-lavender:#b8a4ed;
    --color-brand-peach:#ffb084;
    --color-brand-ochre:#e8b94a;
    --color-brand-mint:#a4d4c5;
    --color-success:#22c55e;
    --color-warning:#f59e0b;
    --color-error:#ef4444;
    --radius-xs:6px;
    --radius-sm:8px;
    --radius-md:12px;
    --radius-lg:16px;
    --radius-xl:24px;
    --radius-pill:9999px;
    --sp-xxs:4px;
    --sp-xs:8px;
    --sp-sm:12px;
    --sp-md:16px;
    --sp-lg:24px;
    --sp-xl:32px;
    --sp-xxl:48px;
    --sp-section:96px;
    --font-display:"Inter",sans-serif;
    --font-body:"Inter",sans-serif;
    --max-width:1280px;
}

html{scroll-behavior:smooth;-webkit-text-size-adjust:100%}
body{background:var(--color-canvas);color:var(--color-body);font-family:var(--font-body);font-size:16px;line-height:1.55;-webkit-font-smoothing:antialiased}

a{color:var(--color-ink);text-decoration:underline;text-underline-offset:3px}
a:hover{opacity:.75}
img{max-width:100%;height:auto;display:block}

.container{max-width:var(--max-width);margin:0 auto;padding:0 var(--sp-xl)}

.top-nav{background:var(--color-canvas);border-bottom:1px solid var(--color-hairline);position:sticky;top:0;z-index:100;height:64px}
.nav-inner{display:flex;align-items:center;justify-content:space-between;height:64px}
.nav-logo{font-size:18px;font-weight:600;letter-spacing:-.3px;color:var(--color-ink);text-decoration:none;flex-shrink:0}
.nav-logo:hover{opacity:.75}
.nav-links{display:flex;list-style:none;gap:var(--sp-xl);align-items:center}
.nav-link{font-size:14px;font-weight:500;color:var(--color-ink);text-decoration:none;letter-spacing:0;line-height:1.4}
.nav-link:hover{opacity:.65}
.nav-link[aria-current="page"]{color:var(--color-primary);text-decoration:underline;text-underline-offset:4px}
.nav-toggle{display:none;flex-direction:column;gap:5px;background:none;border:none;cursor:pointer;padding:var(--sp-xs)}
.nav-toggle span{display:block;width:22px;height:2px;background:var(--color-ink);border-radius:2px;transition:transform .2s,opacity .2s}

.hero-band{background:var(--color-canvas);padding:var(--sp-section) 0}
.hero-inner{display:grid;grid-template-columns:7fr 5fr;gap:var(--sp-xl);align-items:center}
.hero-tag{display:inline-block;font-size:12px;font-weight:600;letter-spacing:1.5px;text-transform:uppercase;color:var(--color-muted);margin-bottom:var(--sp-md)}
.hero-title{font-size:56px;font-weight:500;line-height:1.05;letter-spacing:-2px;color:var(--color-ink);margin-bottom:var(--sp-lg)}
.hero-sub{font-size:18px;font-weight:400;line-height:1.55;color:var(--color-body-strong);max-width:480px;margin-bottom:var(--sp-xl)}
.hero-image-card{background:var(--color-surface-soft);border-radius:var(--radius-xl);overflow:hidden;aspect-ratio:4/3}
.hero-image-card img{width:100%;height:100%;object-fit:cover}

.btn-primary{display:inline-flex;align-items:center;justify-content:center;background:var(--color-primary);color:var(--color-on-primary);font-size:14px;font-weight:600;line-height:1;padding:12px 20px;height:44px;border-radius:var(--radius-md);text-decoration:none;border:none;cursor:pointer;transition:background .15s}
.btn-primary:hover{background:var(--color-primary-active);opacity:1}
.btn-secondary{display:inline-flex;align-items:center;justify-content:center;background:var(--color-canvas);color:var(--color-ink);font-size:14px;font-weight:600;line-height:1;padding:12px 20px;height:44px;border-radius:var(--radius-md);text-decoration:none;border:1px solid var(--color-hairline);cursor:pointer;transition:border-color .15s}
.btn-secondary:hover{border-color:var(--color-ink);opacity:1}
.btn-on-color{display:inline-flex;align-items:center;justify-content:center;background:var(--color-canvas);color:var(--color-ink);font-size:14px;font-weight:600;padding:12px 20px;height:44px;border-radius:var(--radius-md);text-decoration:none;border:none;cursor:pointer}

section{padding:var(--sp-section) 0}
.section-label{font-size:12px;font-weight:600;letter-spacing:1.5px;text-transform:uppercase;color:var(--color-muted);margin-bottom:var(--sp-md)}
.section-title{font-size:40px;font-weight:500;line-height:1.1;letter-spacing:-1px;color:var(--color-ink);margin-bottom:var(--sp-lg)}
.section-sub{font-size:16px;color:var(--color-body);max-width:600px;margin-bottom:var(--sp-xxl)}

.feature-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:var(--sp-lg)}
.feature-card{border-radius:var(--radius-xl);padding:var(--sp-xl)}
.feature-card-pink{background:var(--color-brand-pink);color:var(--color-on-primary)}
.feature-card-teal{background:var(--color-brand-teal);color:var(--color-on-dark)}
.feature-card-lavender{background:var(--color-brand-lavender);color:var(--color-ink)}
.feature-card-peach{background:var(--color-brand-peach);color:var(--color-ink)}
.feature-card-ochre{background:var(--color-brand-ochre);color:var(--color-ink)}
.feature-card-cream{background:var(--color-surface-card);color:var(--color-ink)}
.feature-card h3{font-size:18px;font-weight:600;line-height:1.4;margin-bottom:var(--sp-sm)}
.feature-card p{font-size:14px;line-height:1.55;opacity:.85}
.feature-card .card-icon{font-size:28px;margin-bottom:var(--sp-md)}
.feature-card img{border-radius:var(--radius-md);margin-bottom:var(--sp-md);width:100%;height:160px;object-fit:cover}

.articles-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:var(--sp-lg)}
.article-card{background:var(--color-canvas);border:1px solid var(--color-hairline);border-radius:var(--radius-lg);overflow:hidden;text-decoration:none;color:inherit;transition:border-color .15s,box-shadow .15s;display:flex;flex-direction:column}
.article-card:hover{border-color:var(--color-ink);box-shadow:0 4px 16px rgba(10,10,10,.06);opacity:1}
.article-card-img{aspect-ratio:16/9;overflow:hidden}
.article-card-img img{width:100%;height:100%;object-fit:cover;transition:transform .3s}
.article-card:hover .article-card-img img{transform:scale(1.03)}
.article-card-body{padding:var(--sp-lg);flex:1;display:flex;flex-direction:column}
.article-tag{display:inline-block;font-size:12px;font-weight:600;letter-spacing:1.5px;text-transform:uppercase;color:var(--color-muted);margin-bottom:var(--sp-xs)}
.article-card h3{font-size:18px;font-weight:600;line-height:1.4;letter-spacing:-.3px;color:var(--color-ink);margin-bottom:var(--sp-sm);flex:1}
.article-card p{font-size:14px;color:var(--color-muted);line-height:1.55}
.article-date{font-size:13px;font-weight:500;color:var(--color-muted-soft);margin-top:var(--sp-md)}

.bg-surface-soft{background:var(--color-surface-soft)}
.bg-surface-card{background:var(--color-surface-card)}
.bg-surface-dark{background:var(--color-surface-dark)}

.species-grid{display:grid;grid-template-columns:repeat(2,1fr);gap:var(--sp-lg)}
.species-card{background:var(--color-canvas);border:1px solid var(--color-hairline);border-radius:var(--radius-lg);overflow:hidden;display:flex;gap:var(--sp-lg);align-items:flex-start;padding:var(--sp-lg)}
.species-card img{width:120px;height:90px;object-fit:cover;border-radius:var(--radius-md);flex-shrink:0}
.species-card-body h3{font-size:16px;font-weight:600;color:var(--color-ink);margin-bottom:var(--sp-xs)}
.species-card-body .species-latin{font-size:13px;font-style:italic;color:var(--color-muted);margin-bottom:var(--sp-sm)}
.species-card-body p{font-size:14px;color:var(--color-body);line-height:1.55}

.contact-band{background:var(--color-surface-soft);padding:var(--sp-section) 0}
.contact-inner{display:grid;grid-template-columns:1fr 1fr;gap:var(--sp-xxl);align-items:start}
.contact-text h2{font-size:40px;font-weight:500;line-height:1.1;letter-spacing:-1px;color:var(--color-ink);margin-bottom:var(--sp-lg)}
.contact-text p{font-size:16px;color:var(--color-body);line-height:1.55}
.contact-form{background:var(--color-canvas);border-radius:var(--radius-xl);padding:var(--sp-xl);border:1px solid var(--color-hairline)}
.form-group{margin-bottom:var(--sp-lg)}
.form-group label{display:block;font-size:14px;font-weight:600;color:var(--color-body-strong);margin-bottom:var(--sp-xs)}
.form-group input,.form-group textarea{width:100%;background:var(--color-canvas);color:var(--color-ink);font-size:16px;font-family:var(--font-body);padding:12px 16px;height:44px;border:1px solid var(--color-hairline);border-radius:var(--radius-md);outline:none;transition:border-color .15s}
.form-group textarea{height:auto;min-height:110px;resize:vertical}
.form-group input:focus,.form-group textarea:focus{border-color:var(--color-ink)}
.form-submit{width:100%}

.breadcrumb{font-size:13px;font-weight:500;color:var(--color-muted);margin-bottom:var(--sp-xl);display:flex;gap:var(--sp-xs);align-items:center;flex-wrap:wrap}
.breadcrumb a{color:var(--color-muted);text-decoration:none}
.breadcrumb a:hover{color:var(--color-ink)}
.breadcrumb-sep{color:var(--color-muted-soft)}

.article-header{padding:var(--sp-section) 0 var(--sp-xxl)}
.article-header h1{font-size:48px;font-weight:500;line-height:1.08;letter-spacing:-1.5px;color:var(--color-ink);max-width:820px;margin-bottom:var(--sp-lg)}
.article-meta{display:flex;gap:var(--sp-lg);align-items:center;flex-wrap:wrap;margin-bottom:var(--sp-xl)}
.article-meta .meta-item{font-size:14px;color:var(--color-muted)}
.article-hero-img{border-radius:var(--radius-xl);overflow:hidden;aspect-ratio:21/9;margin-bottom:var(--sp-xxl)}
.article-hero-img img{width:100%;height:100%;object-fit:cover}
.article-body{display:grid;grid-template-columns:1fr 300px;gap:var(--sp-xxl);align-items:start}
.article-content{max-width:720px}
.article-content h2{font-size:32px;font-weight:500;line-height:1.15;letter-spacing:-.5px;color:var(--color-ink);margin:var(--sp-xxl) 0 var(--sp-lg)}
.article-content h3{font-size:22px;font-weight:600;line-height:1.3;color:var(--color-ink);margin:var(--sp-xl) 0 var(--sp-md)}
.article-content p{font-size:16px;color:var(--color-body);line-height:1.65;margin-bottom:var(--sp-lg)}
.article-content ul,.article-content ol{padding-left:var(--sp-xl);margin-bottom:var(--sp-lg)}
.article-content li{font-size:16px;color:var(--color-body);line-height:1.65;margin-bottom:var(--sp-xs)}
.article-content a{color:var(--color-ink);font-weight:500}
.article-content strong{color:var(--color-body-strong);font-weight:600}
.article-content .callout{background:var(--color-surface-card);border-radius:var(--radius-lg);padding:var(--sp-lg) var(--sp-xl);margin:var(--sp-xl) 0;border-left:4px solid var(--color-brand-ochre)}
.article-content .callout p{margin:0}
.article-sidebar{position:sticky;top:80px}
.sidebar-card{background:var(--color-surface-card);border-radius:var(--radius-lg);padding:var(--sp-lg);margin-bottom:var(--sp-lg)}
.sidebar-card h4{font-size:14px;font-weight:600;letter-spacing:1.5px;text-transform:uppercase;color:var(--color-muted);margin-bottom:var(--sp-md)}
.sidebar-card ul{list-style:none;display:flex;flex-direction:column;gap:var(--sp-sm)}
.sidebar-card ul li a{font-size:14px;font-weight:500;color:var(--color-ink);text-decoration:none;line-height:1.4}
.sidebar-card ul li a:hover{text-decoration:underline}

.related-articles{background:var(--color-surface-soft);padding:var(--sp-section) 0}

.page-hero{padding:var(--sp-section) 0 var(--sp-xxl)}
.page-hero h1{font-size:48px;font-weight:500;line-height:1.08;letter-spacing:-1.5px;color:var(--color-ink);margin-bottom:var(--sp-lg)}
.page-hero p{font-size:18px;color:var(--color-body-strong);max-width:600px}
.page-content{max-width:720px;padding-bottom:var(--sp-section)}
.page-content h2{font-size:28px;font-weight:600;line-height:1.2;color:var(--color-ink);margin:var(--sp-xxl) 0 var(--sp-md)}
.page-content p{font-size:16px;color:var(--color-body);line-height:1.65;margin-bottom:var(--sp-lg)}
.page-content ul{padding-left:var(--sp-xl);margin-bottom:var(--sp-lg)}
.page-content li{font-size:16px;color:var(--color-body);line-height:1.65;margin-bottom:var(--sp-xs)}
.page-content a{color:var(--color-ink);font-weight:500}
.page-content address{font-style:normal}

.badge-pill{display:inline-flex;align-items:center;background:var(--color-surface-card);color:var(--color-ink);font-size:13px;font-weight:500;padding:4px 12px;border-radius:var(--radius-pill)}

.cookie-banner{position:fixed;bottom:0;left:0;right:0;background:var(--color-surface-dark);color:var(--color-on-dark);padding:var(--sp-lg) var(--sp-xl);z-index:1000;display:flex;align-items:center;justify-content:space-between;gap:var(--sp-xl);flex-wrap:wrap}
.cookie-banner p{font-size:14px;color:var(--color-on-dark);line-height:1.55;max-width:640px}
.cookie-banner p a{color:var(--color-brand-mint)}
.cookie-actions{display:flex;gap:var(--sp-sm);flex-shrink:0}
.cookie-accept{background:var(--color-canvas);color:var(--color-ink);font-size:14px;font-weight:600;padding:10px 20px;border-radius:var(--radius-md);border:none;cursor:pointer}
.cookie-reject{background:transparent;color:var(--color-on-dark);font-size:14px;font-weight:600;padding:10px 20px;border-radius:var(--radius-md);border:1px solid rgba(255,255,255,.25);cursor:pointer}
.cookie-reject:hover{border-color:rgba(255,255,255,.5)}

.updated-label{font-size:13px;font-weight:500;color:var(--color-muted-soft);margin-bottom:var(--sp-lg);display:block}

@media(max-width:1024px){
    .feature-grid{grid-template-columns:repeat(2,1fr)}
    .articles-grid{grid-template-columns:repeat(2,1fr)}
    .hero-title{font-size:44px}
    .section-title{font-size:34px}
}

@media(max-width:768px){
    .container{padding:0 var(--sp-lg)}
    .nav-links{display:none;position:absolute;top:64px;left:0;right:0;background:var(--color-canvas);border-bottom:1px solid var(--color-hairline);flex-direction:column;gap:0;padding:var(--sp-md) 0}
    .nav-links.is-open{display:flex}
    .nav-links li a{display:block;padding:var(--sp-sm) var(--sp-xl)}
    .nav-toggle{display:flex}
    .hero-inner{grid-template-columns:1fr}
    .hero-title{font-size:36px;letter-spacing:-1px}
    .hero-image-card{display:none}
    .feature-grid{grid-template-columns:1fr}
    .articles-grid{grid-template-columns:1fr}
    .species-grid{grid-template-columns:1fr}
    .contact-inner{grid-template-columns:1fr}
    .article-body{grid-template-columns:1fr}
    .article-sidebar{display:none}
    .article-header h1{font-size:32px}
    .page-hero h1{font-size:32px}
    .section-title{font-size:28px}
    .cookie-banner{flex-direction:column;align-items:flex-start}
    .hero-band{padding:var(--sp-xxl) 0}
    section{padding:var(--sp-xxl) 0}
}